bhajyan madhyaiś cala-kuca-paṭaiḥ kuṇḍalair gaṇḍa-lolaiḥ
svidyan-mukhyaḥ kavara-rasanāgranthayaḥ kṛṣṇa-vadhvo
gāyantyas taṁ taḍita iva tā megha-cakre virejuḥ
Śrīla Śrīdhara Svāmī explains that according to the analogy of lightning flashing in clouds, the perspiration on the lovely faces of the gopīs resembled drops of mist, and their singing resembled thunder. The word āgranthayaḥ may also be read agranthayaḥ, meaning “loosened.” This would indicate that although the gopīs began the dance with their hair and belts tightly drawn, these gradually slackened and loosened.
Śrīla Viśvanātha Cakravartī points out that the gopīs were expert at exhibiting mudrās (precise hand gestures that express feelings or convey meanings associated with the theme of a performance). Thus sometimes Kṛṣṇa and the gopīs would artistically move their interlocked arms together, and sometimes they would separate arms and exhibit mudrās to act out the meaning of the songs they were singing.
The word pāda-nyāsaiḥ indicates that the gopīs artistically and gracefully placed the steps of their dancing feet in an enchanting way, and the words sa-smitair bhrū-vilāsair indicate that the romantic movements of their eyebrows, smiling with love, were most charming to behold.